
1793 illustration of the quagga stallion of Louis XVI's menagerie at Versailles.

Quagga specimen at Natural History Museum, London.
The 'quagga' is an
extinct subspecies of the
plains zebra, which was once found in great numbers in
South Africa's
Cape Province and the southern part of the
Orange Free State. It was distinguished from other zebras by having the usual vivid marks on the front part of the body only. In the mid-section, the stripes faded and the dark, inter-stripe spaces became wider, and the hindquarters were a plain brown. The name comes from a
Khoikhoi word for ''zebra'' and is
onomatopoeic, being said to resemble the quagga's call.
Taxonomy
The quagga was originally classified as an individual
species, ''Equus quagga'', in 1788. Over the next fifty years or so, many other zebras were described by naturalists and explorers. Because of the great variation in coat patterns (no two zebras are alike), taxonomists were left with a great number of described "species", and no easy way to tell which of these were true species, which were
subspecies, and which were simply natural variants.
Long before this confusion was sorted out, the quagga had been hunted to extinction for meat, hides, and to preserve feed for domesticated stock. The last wild quagga was probably shot in the late 1870s, and the last specimen in captivity died on
August 12,
1883 at the
Artis Magistra zoo in
Amsterdam. Because of the great confusion between different zebra species, particularly among the general public, the quagga had become extinct before it was realized that it appeared to be a separate species.
The quagga was the first extinct creature to have its
DNA studied. Recent genetic research at the
Smithsonian Institution has demonstrated that the quagga was in fact not a separate species at all, but diverged from the extremely variable
plains zebra, ''Equus burchelli'', between 120,000 and 290,000 years ago, and suggests that it should be named ''Equus burchelli quagga''. However, according to the rules of
biological nomenclature, where there are two or more alternative names for a single species, the name first used takes priority. As the quagga was described about thirty years earlier than the plains zebra, it appears that the correct terms are ''E. quagga quagga'' for the quagga and ''E. quagga burchelli'' for the plains zebra.
After the very close relationship between the quagga and surviving zebras was discovered, the
Quagga Project was started by
Reinhold Rau in South Africa to recreate the quagga by selective breeding from plains zebra stock, with the eventual aim of reintroducing them to the wild. This type of breeding is also called ''
breeding back''. In early 2006, it was reported that the third and fourth generations of the project have produced animals which look very much like the depictions and preserved specimens of the quagga, though whether looks alone are enough to declare that this project has produced a true "re-creation" of the original quagga is controversial.
DNA from mounted specimens was successfully extracted in 1984, but the technology to use recovered DNA for breeding does not yet exist. In addition to skins such as the one held by the Natural History Museum in London, there are 23 known stuffed and mounted quagga throughout the world. A twenty-fourth specimen was destroyed in
Königsberg, Germany during
World War II.
[1]
Quagga hybrids and similar animals
Zebras have been cross-bred to other equines such as
donkeys and
horses. There are modern animal farms which continue to do so. The offspring are known as
zeedonks,
zonkeys and
zorses (the term for all such zebra hybrids is
zebroid). Zebroids are often exhibited as curiosities although some are broken to harness or as riding animals. On January 20, 2005, Henry, a foal of the Quagga Project, was born. He most resembles the quagga.
There is a record of a quagga being bred to a horse in the 1896 work ''Anomalies and Curiosities of Medicine'' by George M. Gould and Walter L. Pyle:
:''In the year 1815 Lord Morton put a male quagga to a young chestnut mare of seven-eighths Arabian blood, which had never before been bred from. The result was a female hybrid which resembled both parents.''
[2]
In his 1859 ''
The Origin of Species'',
Charles Darwin recalls seeing coloured drawings of zebra-donkey hybrids, and mentions ''"Lord Moreton's famous hybrid from a chesnut'' [sic] ''mare and male quagga..."'' Darwin mentioned this particular hybrid again in 1868 in ''The Variation Of Animals And Plants Under Domestication'',
[3] and provides a citation to the journal in which Lord Morton first described the breeding.
Okapi markings are nearly the reverse of the quagga, with the forequarters being mostly plain and the hindquarters being heavily striped. However, the okapi is no relation of the quagga, horse, donkey, or zebra. Its closest taxonomic relative is the
giraffe.
Trivia
"Quagga" is the code name for the software that runs the
Free Software Foundation Free Software Directory, due to its phonetic awkwardness on par with
GNU.
The
Quagga open source project is a
fork of the
GNU Zebra routing software project. In contrast to the fate of the species, the Quagga software lives on while the Zebra software is largely extinct.
The
Quagga mussel was so named due to its resemblance to the
Zebra mussel. In Lake Michigan, where both are
invasive species, the Quagga mussel has largely out-competed and replaced the more well-known Zebra.
[4]
The alternate ending to
Racing Stripes features a "zorse" (half zebra half horse), which could be a quagga.
See also
★
List of extinct animals
References
1. Max D.T. 2006. Can You Revive an Extinct Animal? The New York Times. Published: January 1 2006. http://www.nytimes.com/2006/01/01/magazine/01taxidermy.html.
2. Hartwell, S. Hybrid Mammals. Downloaded at July 24 2006 at http://www.messybeast.com/genetics/hybrid-mammals.html.
3. Darwin, C. 1883. The Variation of Animals and Plants Under Domestication. Second Edition, Revised. D. Appleton & Co, New York. Online available at http://www.esp.org/books/darwin/variation/facsimile/title3.html.
4. Great Lakes WATER Institute. Downloaded on June 5 2007 from http://www.glwi.uwm.edu/features/news/InWisconsinQuaggas.php.
Other sources
★ , M.A., East, R. & Rubenstein, D.I. 2002. Equus quagga. In:
2006 IUCN Red List of Threatened Species. Last accessed on 08 March 2007
External links
★ ARKive -
images and movies of the quagga ''(Equus quagga)''
★
The Quagga Project
★
PBS Nature: Restoring the Quagga
★
Biology Letters: A rapid loss of stripes: the evolutionary history of the extinct quagga, 2005
★
The Extinction Website - Species Info - Quagga
★
African Explorer - Bringing the Quagga back to life.
★
"Quagga Quest Can we bring back a long-extinct animal?"