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NITRIFYING BACTERIA

'Nitrifying bacteria' are chemoautotrophic bacteria that grow at the expense of inorganic nitrogen compounds.[1] Many species of nitrifying bacteria have complex internal membrane systems that are the location for key enzymes in nitrification: ammonia monooxygenase which oxidizes ammonia to hydroxylamine, and nitrite oxidoreductase, which oxidizes nitrite to nitrate.

Contents
Ecology
Oxidation of ammonia to nitrate
See also
References

Ecology


Nitrifying bacteria are widespread in soil and water, and are found in highest numbers where considerable amounts of ammonia are present (areas with extensive protein decomposition, and sewage treatment plants).[2] Nitrifying bacteria thrive in lakes and streams high imputs of sewage and wastewater because of the high ammonia content.

Oxidation of ammonia to nitrate


Nitrification in nature is the result of actions of two groups of organisms, the 'nitrosifyers' (ammonia-oxidizing bacteria) and 'nitrifying bacteria' (nitrite-oxidizing, nitrate producing bacteria)[3]
'Nitrosifying bacteria'

1. NH3 + O2 + 2e- + 2H+ → NH2OH + H2O

2. NH2OH + H2O + 1/2 O2 → NO2- +2 H2O + H+

'Nitrifying bacteria'

NO2- + 1/2 O2 → NO3-
'Nitrifying bacteria that oxidize ammonia (nitrosifyers)'CharacteristicsGenusPhylogenetic groupDNA (mol% GC)Habitats
Gram-negative short to long rods, motile (polar flagella)or nonmotile; peripheral membrane systems ''Nitrosomonas'' Beta 45-53 Soil, Sewage, freshwater, Marine
Large cocci, motile, vesicular or peripheral membranes ''Nitrosococcus'' Gamma 49-50 Freshwater, Marine
Spirals, motile (peritrichous flagella); no obvious membrane system ''Nitrosospira'' Beta 54 Soil
Pleomorphic, lobular, compartmented cells; motile (peritrichous flagella) ''Nitrosolobus'' Beta 54 Soil

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'Nitrifying bacteria that oxidize nitrite'CharacteristicsGenusPhylogenetic groupDNA (mol% GC)Habitats
Short rods, reproduce by budding, occasionally motile (single subterminal flagella) or non-motile; membrane system arranged as a polar cap ''Nitrobacter'' Alpha 59-62 Soil, Freshwater, Marine
Long, slender rods, nonmotile, no obvious membrane system ''Nitrospina'' Delta 58 Marine
Large Cocci, motile (one or two subterminal flagellum) membrane system randomly arranged in tubes ''Nitrococcus'' Gamma 61 Marine
Helical to vibroid-shaped cells; nonmotile; no internal membranes ''Nitrospira'' ''Nitrospira'' group 50 Marine, Soil

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See also



Denitrification

Nitrification

Nitrogen cycle

Nitrogen deficiency

Nitrogen fixation

References


1. The nature of nitrogen: an overview, Mancinelli RL, , , Life support & biosphere science : international journal of earth space, 1996
2. Population ecology of nitrifying bacteria, Belser LW, , , Annu. Rev. Microbiol., 1979
3. Nitrification and ammonification in aquatic systems, Ward BB, , , Life support & biosphere science : international journal of earth space, 1996


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