The spider family 'Liphistiidae' comprises 5
genera and 87
species [1]. They are among the most
primitive living
spiders, belonging to the suborder
Mesothelae. They are rarely seen tube-dwelling spiders from
Southeast Asia,
China, and
Japan that construct rudimentary trap-doors, characterized by their downward pointing, daggerlike
chelicerae without venom glands
[1], and by having a segmented series of plates on the upper surface of their abdomens. Some make silk trip-lines radiating away from the burrow entrance. They are active at night and live for many years, and females molt aver maturity. Adult males wander in search for females, which rarely leave their burrows. The respiratory system consists only of
booklungs, which could be a reason why these spiders show a quite low level of activity.
[Coddington, J.A. & Levi, H.W. (1991). Systematics and Evolution of Spiders (Araneae). ''Annu. Rev. Ecol. Syst.'' 22:565-592]
In caves in Malaysia,
[2], three different species of ''Liphistius'' are known, and each species is endemic to just one or two caves. The most well known of them is ''
Liphistius batuensis'', which is found in
Batu Caves. In
Japan, the
Kimura-gumo (''Heptathela kimurai'') is rather well-known.
Although they are
Orthognatha (having downward pointing chelicerae), like the
Mygalomorphae, there is no close relationship between those two. It is thought that the common ancestor of all spiders was orthognath, and that in the
Opisthothelae, comprising of Mygalomorphae (mostly tarantulas) and
Araneomorphae (all other spiders), only the Araneomorphae changed their alignment of chelicerae, while the mygalomorphs retained this
symplesiomorph feature.
[Coddington, J.A. & Levi, H.W. (1991). Systematics and Evolution of Spiders (Araneae). ''Annu. Rev. Ecol. Syst.'' 22:565-592]
Genera
★ ''
Heptathela'' Kishida, 1923 (Vietnam, Japan, China)
★ ''
Liphistius'' Schiödte, 1849 (Southeast Asia)
★ ''
Nanthela'' Haupt, 2003 (Hong Kong, Vietnam)
★ ''
Ryuthela'' Haupt, 1983 (
Ryukyu Islands, Okinawa)
★ ''
Songthela'' Ono, 2000 (China)
See also
★
List of Liphistiidae species
★
Spider families
References
1. Haupt, J. (2004). The Mesothelae - a monograph of an exceptional group of spiders (Araneae: Mesothelae). ''Zoologica'' 154:8 , ISBN 3-510-55041-2 (Abstract)
★ Ono, H. (1999) Spiders of the genus ''Heptathela'' (Araneae, Liphistiidae) from Vietnam, with notes on their natural history. The Journal of Arachnology 27(1): 37-43.
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