In
photosynthesis, the 'light-independent reactions', also somewhat misleadingly called the 'dark reactions' (they don't require darkness, but they do require the ''products'' of the
light reactions), are
chemical reactions that convert
carbon dioxide and other
compounds into
glucose. It occurs in the stroma, the fluid filled area of a chloroplast outside of the thylakoid membranes. These reactions, unlike the
light-dependent reactions, do not need
light to occur; hence the term ''dark reactions''. These reactions take the products of the light-dependent reactions and perform further chemical processes on them. There are two light-independent reactions:
carbon fixation and the
Calvin-Benson cycle.
However in
CAM (Crassulacean acid metabolism) plants, carbon fixation actually ''does'' take place at night.
Carbon fixation
''Main article'': ''
carbon fixation''
The
carbon fixation reaction is the first step of the light-independent reactions.
Carbon from
carbon dioxide is "fixed" into a larger
carbohydrate. Three
pathways to occur:
C3 carbon fixation (the most common),
C4 carbon fixation, and CAM (Crassulacean Acid Metabolism). C3 fixation occurs as the first step of the Calvin-Benson cycle in all plants. C4 plants first fix carbon dioxide into
malate, which is then used to supply carbon dioxide in the middle of the night to the Calvin-Benson cycle. CAM plants perform a similar process.
Calvin cycle
The
Calvin-Benson cycle takes
carbon dioxide and converts it to
glucose, which the plant uses for energy.
External links
★
The Biochemistry of the Calvin Cycle at Rensselaer Polytechnic Institute