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'''Dilophosaurus''' was a theropod dinosaur from the Early Jurassic Period. The name is pronounced // ("dy-LOH-fo-sawr-us") meaning 'two-crested lizard', because it had two crests (Greek ''di'' meaning 'two', ''lophos'' meaning 'crest' and ''sauros'' meaning 'lizard')

Contents

Description

Skull and crests

Species

In popular culture

References

Description

''Dilophosaurus'' measured around 6 meters (20 ft) long and may have weighed half a ton. Fossils of the animal have been found at the Navajo Indian Reservation, just west of Tuba City, Arizona, USA. Just a few tens of feet below the level of the bones, large footprints of carnivorous dinosaurs were found and these may belong to ''Dilophosaurus''. It lived in the Early Jurassic Period.
The original description was published in 1954, by the renowned paleontologist Samuel Welles. However, at the time, it was thought to be another genus of theropod (''Megalosaurus''). In 1970, it was recognized to be distinct and given its own generic name ''Dilophosaurus''. Welles later redescribed the entire taxon in 1984, in a more comprehensive paper. ''Dilophosaurus'' may be a primitive member of the clade containing both ceratosaurian and tetanuran theropods. Alternatively, some paleontologists classify this genus as a large coelophysoid. Recent potential ''Dilophosaurus'' 'skin imprints', associated with a set of footprints, seemed to suggest that it had feathers but further study revealed these to be impressions of plant material.
Footprints of a ''Dilophosaurus''-like animal have been discovered in Vallåkra in Sweden, in the Lavini di Marco area, near Rovereto (Trentino, northern Italy), and near Rocky Hill (Connecticut). They are associated with those of a ''Camptosaurus''-like animal.

Skull and crests

The most distinctive characteristic of ''Dilophosaurus'' was the pair of rounded crests on its skull, probably used for display. Studies by Rob Gay (2001) show that these crests may have been larger in one sex than in the other. Another curious skull feature was a notch behind the first row of teeth, giving it an almost crocodile-like appearance. This "notch" existed by virtue of a weak connection between the premaxillary and maxillary bones of the skull. This conformation led to the early hypothesis that ''Dilophosaurus'' scavenged off dead carcasses, with the front teeth being too weak to bring down and hold large prey. A similar notch is present in most other species of coelophysoid.

Species

There is another species of ''Dilophosaurus'' (''D. sinensis''), which may or may not belong to this genus. It is possibly closer to the bizarre Antarctic theropod ''Cryolophosaurus'', based on the fact that the anterior end of the jugal does not participate in the internal antorbital fenestra and that the maxillary tooth row is completely in front of the orbit and ends anterior to the vertical strut of the lacrimal. This species was recovered from the Yunnan Province of China in 1987, with the prosauropod ''Yunnanosaurus'' and later described and named in 1993 by Shaojin Hu.

In popular culture

''Dilophosaurus'' was featured in the 1993 movie ''Jurassic Park'' and in the original novel by Michael Crichton (on which the film was based). In the film version, it had a retractable neck frill around its neck (much like a frill-necked lizard), and was able to spit blinding poison, (much like the spitting cobra) aiming for the eyes to blind and paralyze its prey. There is no evidence to support this representation, which was invented by the author and director to heighten suspense. In the film the director Steven Spielberg reduced its size, from moderately large to about 3 feet tall and 5 feet long. This was explained in a quote in the movie by portraying the dinosaur as a juvenile ("you're not so bad as your big brothers"). In the novel, the creature was 10 feet tall with no frill and had poisonous saliva that was used in predation by biting or spitting at its prey. In the novel, this trait was discovered by the researchers at Jurassic Park, who planned to remove the glands containing the poison. This plan was not carried out - an autopsy would have been needed to see how to remove the glands, but it would have required killing a ''Dilophosaurus'', which John Hammond, owner of the park, opposed.
''Jurassic Park'' merchandise, including toys and video games (such as '' and the arcade games '' and ''Jurassic Park III''), often include ''Dilophosaurus''. Another video game, ''ParaWorld'', features a ''Dilophosaurus'' modelled after the ones in ''Jurassic Park''. Yet another dinosaur-themed game, ''Jurassic Wars'', featured a Dilophosaurus, which spat venomous saliva, just as described in the movie.
''Dilophosaurus'' was also featured in the documentary ''When Dinosaurs Roamed America'', killing an ''Anchisaurus'' and scaring off a pack of ''Megapnosaurus''.
On Married with Children, the dilophosaur is mentioned. In episode 807 (Take My Wife, Please), Al has a dinosaur costume on while answering Trick-Or-Treaters on Halloween. When one boy dressed as a dilophosaur is at the door, he tells Al he is "the one that spits."
On an episode of Family Guy, Peter has a toy idea called Facts of LifeTransformers that features The Facts of Life characters who transform into animals "I got another one where Natalie turns into one of those spittin' lizards from Jurassic Park".

References

★ Evidence for sexual dimorphism in the Early Jurassic theropod dinosaur, ''Dilophosaurus'' and a comparison with other related forms, Gay, R., , , Journal of Vertebrate Paleontology, 2001

★ New specimens of ''Dilophosaurus wetherilli'' (Dinosauria: Theropoda) from the early Jurassic Kayenta Formation of northern Arizona, Gay, R., , , Western Association of Vertebrate Paleontologists annual meeting volume, 2001

★ Hu, Shaojin (1993). "A note on a double-crested dragon (''Dilophosaurus'') fossil discovered in Jinning County, Yunnan Province." ''Gu Jizhui Dongwu Xuebao = Vertebr. PalAsiatica'', '31' i.

★ New Jurassic dinosaur from the Kayenta formation of Arizona, Welles, S. P., , , Bulletin of the Geological Society of America, 1954

★ Welles, S. P. (1984). "''Dilophosaurus wetherilli'' (Dinosauria, Theropoda), osteology and comparisons." ''Palaeontogr. Abt. A: Paläozool., Stratigr.'', 185, 85-180.