The bird family 'Casuariidae' has four surviving members: the three
species of
cassowary, and the only remaining species of
Emu. The emus were formerly classified in their own family, Dromaiidae, but are regarded as sufficiently closely related to the cassowaries to be part of the same family.
All four members of the family are very large flightless birds native to
Australia-New Guinea. The characteristics of the family are those of its members.
Systematics and evolution
The emus form a distinct subfamily, characterized by legs adapted for running. As with all
ratites, there are several contested theories concerning their evolution and relationships. As regards this family, it is especially interesting whether emus or cassowaries are the more primitive form: the latter are generally assumed to retain more
plesiomorphic features, but this does not need to be true at all; the fossil record is also ambiguous, and the present state of
genomics does not allow for suffiently comprehensive analyses. A combination of all these approaches with considerations of
plate tectonics at least is necessary for resolving this issue.
The number of cassowary species described based on minor differences in casque shape and color variations is quite large. In recent times, however, only 3 species are recognized, and most authorities only acknowledge few
subspecies or none at all.
The fossil record of casuariforms is interesting, but not very extensive. Regarding fossil species of ''Dromaius'' and ''Casuarius'', see their genus pages.
Some Australian fossils initially believed to be from emus were recognized to represent a distinct genus, ''Emuarius''
[1],
which had a cassowary-like skull and
femur and an emu-like lower leg and foot. In addition, the first fossils of
mihirungs were initially believed to be from giant emus
[2],
but these birds were completely unrelated.
Subfamily 'Casuariinae' - cassowaries
★ Genus '''Casuarius'''
★
★
Southern Cassowary, ''Casuarius casuarius''
★
★
Dwarf Cassowary, ''Casuarius bennetti''
★
★
Northern Cassowary, ''Casuarius unappendiculatus''
Subfamily 'Dromaiinae' - emus
★ Genus '''Dromaius'''
★
★
Emu, ''Dromaius novaehollandiae''
★
★
★
Tasmanian Emu, ''D. n. diemenensis'' (
extinct)
★
★
★
South Eastern Emu, ''D. n. novaehollandiae''
★
★
★
South Western Emu, ''D. n. rothschildi''
★
★
★
Northern Emu, ''D. n. woodwardi''
★
★
Kangaroo Island Emu, ''Dromaius baudinianus'' (
extinct)
★
★
King Island Emu, ''Dromaius ater'' (
extinct)
★ Genus '''Emuarius''' - "emuwaries" (
fossil)
★
★ ''Emuarius guljaruba'' (Late
Oligocene - Late
Miocene)
★
★ ''Emuarius gidju'' (Wipajiri Early
Miocene of Lake Ngapakaldi)
References
★ Boles, Walter E. (2001): A new emu (Dromaiinae) from the Late Oligocene Etadunna Formation. ''
Emu '101': 317–321.
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Footnotes
1.
From "Emu" + "''Casuarius''". Describer W. E. Boles commonly refers to the genus as "emuwaries" or "cassomus".
2.
The vernacular name "mihirung" is derived from ''mihirung paringmal'', which means "giant emu" in the Chaap Wuurong language