The order 'Anseriformes' contains about 150
species of
birds in three families: the
Anhimidae (the screamers),
Anseranatidae (the
Magpie-goose), and the
Anatidae, which includes over 140 species of
waterfowl, among them the
ducks,
geese, and
swans.
All species in the order are highly adapted for an aquatic existence at the water surface. All are web-footed for efficient swimming (although some have subsequently become mainly terrestrial).
Evolution

Magpie Goose ''Anseranas semipalmata''
The earliest known Anseriform is the recently discovered ''
Vegavis'', which lived during the
Cretaceous period
[1]. It is thought that the Anseriformes originated when the original
Galloanserae (the group to which Anseriformes and
Galliformes belong) split into the two main lineages. The extinct
dromornithids represent early offshoots of the anseriform line, possibly derived from
screamer-like ancestors, and so maybe
Gastornis (if it is an Anseriform). The ancestors of the Anseriformes developed the characteristic bill structure that they still share. The combination of the internal shape of the bill and a modified tongue acts as a suction pump to draw water in at the tip of the bill and expel it from the sides and rear; an array of fine filter plates called ''lamellae'' traps small particles, which are then licked off and swallowed.
All Anseriformes have this basic structure, but many have subsequently adopted alternative feeding strategies: geese graze on plants, the
saw-billed ducks catch fish; even the
screamers, which have bills that seem on first sight more like those of the game birds, still have vestigal lamellae. The
prehistoric wading
presbyornithids and the huge and possibly
carnivorous dromornithids were even more bizarre.
Systematics
The Anseriformes and the
Galliformes (pheasants etc) are the most primitive neognathous birds, and should follow
ratites and
tinamous in bird classification systems.
Anatidae systematics, especially regarding placement of some "odd" genera in the dabbling ducks or shelducks, is not fully resolved. See the Anatidae article for more information, and for alternate taxonomic approaches.
'ORDER ANSERIFORMES'
★ Family
Anhimidae: screamers
★ Family
Anseranatidae: the Magpie-goose
★ Family
Anatidae
★
★ Subfamily
Dendrocygninae: Whistling ducks (sometimes given full family status as the Dendrocygnidae).
★
★ Subfamily
Thalassorninae: the White-backed Duck.
★
★ Subfamily
Anserinae: Swans and geese.
★
★ Subfamily
Stictonettinae: the Freckled Duck.
★
★ Subfamily
Plectropterinae: the Spur-winged Goose.
★
★ Subfamily
Tadorninae: Shelducks and sheldgeese - probably
paraphyletic
★
★ Subfamily
Anatinae: Dabbling ducks and moa-nalos
★
★ Subfamily
Aythyinae: Diving ducks (sometimes included in Anatinae)
★
★ Subfamily
Merginae: eiders, scoters, mergansers and other sea-ducks.
★
★ Subfamily
Oxyurinae: Stiff-tailed ducks and allies.
★ Family
Dromornithidae (
fossil): mihirungs
★ Family
Presbyornithidae (
fossil): several genera of wading-"geese", including the oldest known anseriform, ''
Vegavis''.
Some fossil anseriform taxa not assignable with certainty to a family are:
★ ''
Anatalavis'' (Late Cretaceous/Early Paleocene - Early Eocene) - Anseranatidae or
basal. Includes ''"Telmatornis" rex''.
★ ''
Proherodius'' (London Clay Early Eocene of London, England) - Presbyornithidae?
★ ''
Paranyroca'' (Rosebud Early Miocene of Bennett County, USA) - Anatidae or own family?
In addition, a considerable number of Late Cretaceous and
Paleogene fossils have been described where it is uncertain whether or not they are anseriforms. This is due to the fact that almost all orders of aquatic birds living today either originated or underwent a major
radiation during that time, making it hard to decide whether some waterbird-like bone belongs into this family or is the product of parallel evolution in a different lineage due to adaptive pressures.
★ ''
Apatornis'' (Smoky Hill Chalk Late Cretaceous of Twin Butte Creek, USA)
★ "Presbyornithidae" gen. et sp. indet. (Barun Goyot Late Cretaceous of Udan Sayr, Mongolia) - Presbyornithidae?
★ UCMP 117599 (Hell Creek Late Cretaceous of Bug Creek West, USA)
★ ''
Petropluvialis'' (Late Eocene of England) - may be same as ''Palaeopapia''
★ ''
Agnopterus'' (Late Eocene - Late Oligocene of Europe) - includes ''Cygnopterus lambrechti''
★ ''"Headonornis hantoniensis"'' BMNH PAL 4989 (Hampstead Early Oligocene of Isle of Wight, England) - formerly ''"Ptenornis"''
★ ''
Palaeopapia'' (Hampstead Early Oligocene of Isle of Wight, England)
★ ''"Anas" creccoides'' (Early/Middle Oligocene of Belgium)