'Ankylosauria' is a group of
herbivorous dinosaurs of the order
Ornithischia. It includes the great majority of dinosaurs with
armor in the form of bony scutes, although many of them had other types of armor as well. Ankylosaurs were bulky
quadrupeds, with short, powerful limbs. They are first known to have appeared in the early
Jurassic Period of
China, and persisted until the end of the
Cretaceous Period. They have been found on every continent except
Africa. The first dinosaur ever discovered in
Antarctica was the ankylosaurian ''
Antarctopelta'', fossils of which were recovered from
Ross Island in 1986.
Ankylosauria was first described by
Henry Fairfield Osborn in 1923.
[Obsborn, H. F. (1923). "Two Lower Cretaceous dinosaurs of Mongolia." ''American Museum Novitiates", '95': 1–10.[1]] In the
Linnaean classification system, the group is usually considered a suborder or an infraorder. It is contained within the group
Thyreophora, which also includes the stegosaurians, armored dinosaurs known for their combination of plates and spikes.
Classification
Ankylosauria is split into two
families,
Nodosauridae (the nodosaurids) and
Ankylosauridae (the ankylosaurids). The big difference is that most ankylosaurids (except the "polacanthids") have bony
clubs at the end of their tails, which nodosaurids lack.
The nodosaurids were earlier and more primitive. They had narrow heads, and frequently had large spikes protruding from their bodies. This group traditionally includes ''
Nodosaurus'', ''
Edmontonia'', and ''
Sauropelta''.
The polacanthines, an intermediate group sometimes placed in their own family, were once placed in the Nodosauridae, due to their lack of tail clubs. More recently, it has been found that they were more closely related to the ankylosaurids and, by most
cladistic definitions, are members of that family. The polacanthines include ''
Hylaeosaurus'', ''
Polacanthus'' and ''
Mymoorapelta''. Opinions range from them belonging to their own family, the Polocanthidae,
[1] distinct from the other two or being a subfamily of Ankylosaurs, to some researchers questioning their status as a discrete group.
The traditional ankylosaurids are from later in the
Cretaceous. They had much wider bodies, and thicker armor. They have even been discovered with bony eyelids. The large clubs at the end of their tails were probably used in self-defence, and presumably were swung at predators. This family included ''
Ankylosaurus'', ''
Euoplocephalus'', and ''
Pinacosaurus''.
In 1997,
Carpenter defined the clade Ankylosauria as all thyreophorans closer to ''Ankylosaurus'' than to ''Stegosaurus'' (a definition followed by most paleontologists today, including Sereno, 2005). This "stem-based" definition means that the primitive armored dinosaur ''
Scelidosaurus'', which is slightly closer to ankylosaurids than to stegosaurids, is technically a member of Ankylosauria. Upon the discovery of ''
Bienosaurus'',
Dong Zhiming (2001) erected the family
Scelidosauridae for both of these primitive ankylosaurs.
Taxonomy
A simplified version of one possible classification follows:
★ Order
Ornithischia (the "bird hipped" dinosaurs)
★
★ Suborder
Thyreophora (armored
herbivores)
★
★
★ Infraorder Ankylosauria (the tanks)
★
★
★
★ Family
Scelidosauridae
★
★
★
★
★ ''
Scelidosaurus''
★
★
★
★
★ ''
Bienosaurus''
★
★
★
★ ''
Minmi''
★
★
★
★ ''
Antarctopelta''
★
★
★
★ Family
Nodosauridae
★
★
★
★ Family
Ankylosauridae
★
★
★
★
★ Subfamily Polacanthinae
★
★
★
★
★ Subfamily Ankylosaurinae (tail clubs)
References
1. The Armored Dinosaurs, Carpenter K, , , Indiana University Press, 2001, ISBN 0-253-33964-2
External links
★
Ankylosauria at DinoData